The 1/4-20 holes in the SMW plate are countersunk for use with their Fixturing Pins for alignment (two of these pins are at the bottom of the photo above), which I think are what @GeneT45 was referring to.
No, what I'm referring to is e.g. 3/8"-16 threads use a 5/16" tap drill, so the hole may be used for a 3/8-16 fastener (threaded) or a 5/16 dowel pin without interspersing threaded holes and pin holes. Doesn't look like that synergy exists any smaller than that in Standard sizes, but it might in metric (I didn't check).
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